Thermal fluctuations in Human Cell Nucleus

I recently read an interesting  paper showing  in human cell,   the nucleoli exhibit subtle, but measurable, surface shape fluctuations, consistent with the behavior of a liquid-liquid interface with a low surface tension. As the surface modes follows the rule of energy equipartition theorem, the surface fluctuations should be driven by thermally excited capillary waves. This paper also shows when two nucleus coalesce, the neck connecting two fusing nucleoli grows in time which can be predicted by classic theory without thermal fluctuations, which derives more research since thermal fluctuations are possible to speed up this process (The link of this paper is: